Abstract: Deep learning (DL) rainfall–runoff models outperform conceptual, process-based models in a range of applications. However, it remains unclear whether DL models can produce physically plausible projections of streamflow under climate change. We investigate this question through a sensitivity analysis of modeled responses to increases in temperature and potential evapotranspiration (PET), with other meteorological variables left unchanged. Previous research has shown that temperature-based PET methods overestimate evaporative water loss under warming compared with energy budget-based PET methods. We therefore assume that reliable streamflow responses to warming should exhibit less evaporative water loss when forced with smaller, energy-budget-based PET compared with temperature-based PET. We conduct this assessment using three conceptual, process-based rainfall–runoff models and three DL models, trained and tested across 212 watersheds in the Great Lakes basin. The DL models include a Long Short-Term Memory network (LSTM), a mass-conserving LSTM (MC-LSTM), and a novel variant of the MC-LSTM that also respects the relationship between PET and evaporative water loss (MC-LSTM-PET). After validating models against historical streamflow and actual evapotranspiration, we force all models with scenarios of warming, historical precipitation, and both temperature-based (Hamon) and energy-budget-based (Priestley–Taylor) PET, and compare their responses in long-term mean daily flow, low flows, high flows, and seasonal streamflow timing. We also explore similar responses using a national LSTM fit to 531 watersheds across the United States to assess how the inclusion of a larger and more diverse set of basins influences signals of hydrological response under warming. The main results of this study are as follows: The three Great Lakes DL models substantially outperform all process-based models in streamflow estimation. The MC-LSTM-PET also matches the best process-based models and outperforms the MC-LSTM in estimating actual ...
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